Chasing OG Definitions and Epiphytes  
  

== 2 of 6 ==
Date: Mon, Nov 12 2007 8:51 am
From: dbhguru


Lee,

As I read BVP's and Preston's books, and others that relate results of canopy research in the Pacific Northwest and the Tropics, it become crystal clear at how pathetically little the schools of forestry have heretofore known about old growth ecosystems and what they truly represent. And now, the idea of mimicking or managing for a few old growth characteristics as a surrogate for the real thing in order to provide habitat for several extra species of birds becomes increasingly foolish. In fact down right amateurish. As researchers have discovered, the number of species that inhabit the canopies of the great Pacific Coast rainforests is literally mindboggling. There is not counterpart in younger forests. The plant communities in the OG canopies may endure for the better part of 1,000 years.

With all due respect to my good friend Don Bertolette, I've never bought into the old growth definition of Oliver and Larson. In my view, they see everything from a ground level, timber perspective - verey, very limited. The true measure of old growth, at least in the Pacific Coast forests, may well be measured by the existence of a very wide range of indicator plant and animal communities that inhabit the crowns of loder trees and make the canopy the richest zone of life in old growth forests. I will bet the temperate rainforest and near rainforest areas of the southern Appalachians have a counterpart. Will Blozan has frequently marveled at the lichen encrustations of the huge hemlocks he climbs. The counterpart may be on a much smaller scale, but I'd bet dollars to donuts that it exists.

Whether there is a counterpart in the old growth forests of the Northeast, I don't know. I kind of doubt that one exists, but if it does, I would expect it to exist in the conifers. Anyway, it is interesting to speculate on why there might or might not be a counterpart. Any thoughts on the subject?

Bob


== 3 of 6 ==
Date: Mon, Nov 12 2007 10:34 am
From: DON BERTOLETTE



Bob-
The study of canopies and their associated communities is a 'new science'...efforts in Washington Oregon and northern California (names like Jerry Franklin, Bob Van Pelt, Roman Dial, Steve Sillet) have been underway for several decades, but good science takes awhile...check out Oregon State and UW forestry/ecology curriculums.

As to Oliver and Larson, I'm not especially in love with them, but I do agree that the one absolutely essential ingredient in old-growth ecosystems is time. Were Oliver and Larson being capricious in selecting the amount of time it takes for the cohort that follows the generation of dominant trees responding to a major disturbance...no, I think not.

Is that sufficient time for the epiphytic plant community/upper canopy ecosystem to establish itself in its own old-growth habit?

I don't know, but it seems like a good place to start...what say the members of the forum with upper canopy ecosystem knowledge?

I would bet that the Wind River group (Jerry Franklin and others) is looking closely at this very topic!

-Don



== 4 of 6 ==
Date: Mon, Nov 12 2007 2:47 pm
From: Lee Frelich


Bob:

There probably isn't an exact counterpart in the northeast--too much wind,
fire, and species that don't live as long. Nevertheless, hemlock, white
cedar, and white pine in the northeast might have some canopy communities
that are more complex than in younger forests, but not quite what they have
out west.

Lee



== 5 of 6 ==
Date: Mon, Nov 12 2007 3:44 pm
From: dbhguru


Lee,

BVP points out an interesting fact about shade tolerance that I had not realized and that is that the shade intolerant pines cannot orient their needles to receive maximum solar reception, whereas the shade tolerant firs can. The ability of some conifers to either protect their needles from too much sun or maximize their exposure in the dim light of the forest floor is pretty nifty. I wonder why pines did not develop this neat adaptation.Of course the same thing could be asked about any pioneer species.

Bob


== 6 of 6 ==
Date: Mon, Nov 12 2007 4:02 pm
From: Randy Brown


Bob,

I uploaded a close up picture of a lichen encrusted twig I picked up
along the Boogerman trail in the Smokies this spring:

twig2.jpg (67287 bytes)

Perhaps not Pacific Northwest grade but still an amazing little forest
down in there.


==============================================================================
TOPIC: BVP's Great Book
http://groups.google.com/group/entstrees/browse_thread/thread/c9657775309f4d07?hl=en
==============================================================================

== 1 of 2 ==
Date: Tues, Nov 13 2007 6:03 am
From: "Jess Riddle"


Hi Randy,

Looks like Platismatia tuckermanii on either end of the twig, and you
can see some Pseudevernia consocians above your palm. I'm not sure
what the others are, but it looks like they're providing lots of nice
invertebrate habitat. Both Platismatia and Psuedevernia are common on
old conifers around the Cataloochee section of the Smokies.

Jess


== 2 of 2 ==
Date: Tues, Nov 13 2007 11:19 am
From: DON BERTOLETTE

Bob-
After reading your post below, I thought back to the time that David Kittredge, you, and I went to the Deerfield River Gorge and walked into an old-growth stand you were familiar with. We talked about bark characteristics and gnarly branches, and Dave said "You know, it's like one of the Supreme Court justices said..."I don't know how to define pornography, but I know it when I see it", and Dave went on to say, "I know old-growth when I'm in it, but I'm not sure exactly what is different when I'm out of it", and then he challenged us to more fully identify what 'old-growth' was.
That and other considerations I think spurred both of us on, to more completely define old-growth. Western forest ecologists had a head start, with the pressure of commercial interests wanting to consume them (old-growth forests!). Old-growth forests in the East had been more or less written off, as having already been logged, blown down, herbivoried, or otherwise disturbed.

Reading your post reminded me of how much effort was applied in chasing definitions. It was definitely constructive, it drove us all to further comprehension, but it seemed the closer we got, the less we could definitively say. Your post below pretty much takes us back to 'ground zero'...not a bad place to start! It was several years as I recall, before the medium in which old-growth grows was getting much investigation...soil microorganisms had to be an 'obligate' part of the ecosystem equation, as it didn't take much disturbance of 'ground zero' before the vigor and resilience was affected.

Another consideration was the history (reference conditions) of the stand/forest...what was the natural disturbance regime, had there been significant human entries into the ecosystem, whether is was herbivory imbalance (wildlife consumption outside of natural ranges of variability), domestic grazing, prior indigenous cultural practices, selective logging (winter removals of specific market driven species), or commercially intensive logging.Clearly, in the East, with several centuries of European settlement and millenia of prior indigenous cultural occupation, a restrictive, exclusive definition would preclude what old-growth we walk into and know that it was different before we walked into it.

There is it seems, enough room for elasticity in the old-growth definitions. In a biological context, that elasticity in my mind is the old-growth 'forest health', its 'resilience reserves'...and how much disturbance it absorbs, yet still rebounds in what we might call a natural range of variation over time.

Your comments below about the nature of epiphytic plant communities and that relationship with 'old-growthedness', points out that some of the most important facets of old-growth forests, are not the trees themselves, but the communities that their structures permit. While heterogenous vertical and horizontal structures and species compositions are big words describing tree relationships, they go along way towards providing the basis for the biological diversity that we've come to associate with walking into and out of old-growth forests. At least one more big word should be in above sentence...that word would describe the variations due to time...it takes time before a diverse multi-aged structure evolves, before a varied mosaic of plant composition takes hold, and reflects the reserves necessary for responding to the disturbances it has, and may in the future encounter.
And yes, that is likely to take more than the time it takes for the second cohort following first generation disturbance respondents...

Back atcha!
-Don


=============================================================================
TOPIC: BVP's Great Book
http://groups.google.com/group/entstrees/browse_thread/thread/c9657775309f4d07?hl=en
==============================================================================
== 2 of 2 ==
Date: Tues, Nov 13 2007 11:19 am
From: DON BERTOLETTE

Bob-
After reading your post below, I thought back to the time that David Kittredge, you, and I went to the Deerfield River Gorge and walked into an old-growth stand you were familiar with. We talked about bark characteristics and gnarly branches, and Dave said "You know, it's like one of the Supreme Court justices said..."I don't know how to define pornography, but I know it when I see it", and Dave went on to say, "I know old-growth when I'm in it, but I'm not sure exactly what is different when I'm out of it", and then he challenged us to more fully identify what 'old-growth' was.
That and other considerations I think spurred both of us on, to more completely define old-growth. Western forest ecologists had a head start, with the pressure of commercial interests wanting to consume them (old-growth forests!). Old-growth forests in the East had been more or less written off, as having already been logged, blown down, herbivoried, or otherwise disturbed.

Reading your post reminded me of how much effort was applied in chasing definitions. It was definitely constructive, it drove us all to further comprehension, but it seemed the closer we got, the less we could definitively say. Your post below pretty much takes us back to 'ground zero'...not a bad place to start! It was several years as I recall, before the medium in which old-growth grows was getting much investigation...soil microorganisms had to be an 'obligate' part of the ecosystem equation, as it didn't take much disturbance of 'ground zero' before the vigor and resilience was affected.

Another consideration was the history (reference conditions) of the stand/forest...what was the natural disturbance regime, had there been significant human entries into the ecosystem, whether is was herbivory imbalance (wildlife consumption outside of natural ranges of variability), domestic grazing, prior indigenous cultural practices, selective logging (winter removals of specific market driven species), or commercially intensive logging.Clearly, in the East, with several centuries of European settlement and millenia of prior indigenous cultural occupation, a restrictive, exclusive definition would preclude what old-growth we walk into and know that it was different before we walked into it.

There is it seems, enough room for elasticity in the old-growth definitions. In a biological context, that elasticity in my mind is the old-growth 'forest health', its 'resilience reserves'...and how much disturbance it absorbs, yet still rebounds in what we might call a natural range of variation over time.

Your comments below about the nature of epiphytic plant communities and that relationship with 'old-growthedness', points out that some of the most important facets of old-growth forests, are not the trees themselves, but the communities that their structures permit. While heterogenous vertical and horizontal structures and species compositions are big words describing tree relationships, they go along way towards providing the basis for the biological diversity that we've come to associate with walking into and out of old-growth forests. At least one more big word should be in above sentence...that word would describe the variations due to time...it takes time before a diverse multi-aged structure evolves, before a varied mosaic of plant composition takes hold, and reflects the reserves necessary for responding to the disturbances it has, and may in the future encounter.
And yes, that is likely to take more than the time it takes for the second cohort following first generation disturbance respondents...

Back atcha!
-Don