Monroe State Forest:  Dunbar Brook and Parsonage Brook   Robert Leverett
  Sep 30, 2004 06:49 PDT 

Lee:

   ...On another topic, I'm wondering if you would be willing to give a
quick summary of the forests you saw this past Monday in Monroe State
Forest along Dunbar and Parsonnage Brooks. For the Ents out there who
want to attend the Oct 22nd and 24th event in Monroe State Forest, a
preview of what they would see from a true expert would be highly
valuable. I have tended to over-focus on the individual trees of Dunbar
because until you gave your interpretation, I was pretty confused about
what I was looking at on both the north facing ridge and along
Parsonnage Brook (the last place we visited).

Bob
Re: Request of Lee   Lee E. Frelich
  Sep 30, 2004 09:45 PDT 

Bob:

Yes, here is my report on the two forests.


Dunbar Brook:
Dunbar brook has a large primary forest remnant on steep slopes with many
large boulders, dominated by sugar maple, yellow birch, and white ash, with
some beech, hemlock, red maple, basswood and white pine. The forest
experienced some major blowdown probably about 30 years ago (an exact date
can be determined by coring some of the pole-sized trees that survived and
will show a release from suppression). The blow down was most severe at
mid slope, indicating that the wind hit the slope at an acute angle (i.e.
it was neither parallel to nor at right angles to the valley and adjacent
ridge). This produced turbulence at mid slope and patches of blowdown of
varying size from a few trees to nearly an acre. The heavily hit stands
have an abundance of young synchronously-released trees (total age of these
trees may vary by a century or more because they were suppressed for
varying amounts of time before the storm), with scattered larger trees that
survived the storm. These larger trees are more abundant in boulder-filled
draws that run perpendicular to the ridge, where they had better anchorage
and where slightly less exposed, being rooted 5-10 feet lower than
surrounding trees. Many of the large survivor trees show evidence of branch
and crown breakage from the storm. The developmental stage of these stands
has been variously described as a mature-sapling mosaic, multi-aged pole
stand or disturbed stand with mature remnants. Some stands were not hit as
hard by the wind and have a larger component of large trees with classic
multi-aged forest stage of development. Most of the original northern
hardwood forests from Minnesota to New Hampshire went back and forth from
mature sapling mosaics to old multi-aged stands in response to wind storms
of varying intensity. Therefore this stand provides an ideal exhibit of how
northern hardwoods respond to the type of natural disturbance that is
thought to have perpetuated northern hardwoods for the last several
thousand years.

The successional status of the stands varies little, since the released
understory reflects the late-successional composition of the overstory.
Species that are mid-tolerant of shade, such as white ash, yellow birch,
and red maple, are common in old multi-aged stands because some of the gaps
there are large enough and have enough light to support growth of those
species, even if stands are not hit by storms such as the one that hit
Dunbar Brook a few decades ago. Therefore, primary forest remnants are a
mixture of shade-tolerant and mid-tolerant species, and this mixture forms
the so-called climax species composition. There is a rich moss flora and a
lot of species of wildflowers including nodding trillium.

Hemlock is starting to form a neighborhood fairly high up on the slope,
where there is a cluster of several large trees that vary in age, with
multi-aged saplings surrounding them. Once this patch becomes consolidated,
it could maintain itself until the climate changes. I envision it as
similar to the initial phase of hemlock patch development in Sylvania, MI
that occurred 3000 years ago.

Parsonnage Brook:
On Parsonnage Brook, there is a much different type of primary forest
dominated by hemlock and red spruce, that originated after a severe
stand-levelling disturbance, probably wind, during the mid to late
1800s. As with Dunbar Brook, the canopy has synchronous release but the
total tree age varies because suppressed trees released by the storm were
various ages. What is different here with respect to the disturbance, is
that it was much more intense, and nearly levelled the stand, leaving only
a handful of mature survivors. The stand was probably dominated by
sapling-sized trees at the time the surrounding stands were logged, and is
the likely reason that stand is still in primary condition. At this point,
the first few gaps large enough to recruit new saplings into the canopy are
starting to form (i.e. the stand is reaching the end of the stem exclusion
stage and ready to enter the transition to uneven aged condition).

This stand is dominated by late-successional species now and was right
after the disturbance, at which time it would have been an example of a
young late-successional stand. The disturbance did not cause any
succession, which is defined as directional change in species composition.

This stand has very thick duff, which is a consequence of the high C:N
ratio of the leaf litter of hemlock and red spruce, which is decomposed at
a slow rate, allowing a lot of accumulation. There may be remnants of logs
from the previous canopy buried in the duff, now in category 5 of decay.

This stand has analogs in the highlands of the Porcupine Mountains, MI,
where there are a few hemlock stands (mixed with white spruce and white
cedar in that case) surrounded by hardwoods.

Lee